Human Motor Cortex Reference Explorer

Annotation Markers

class

Label	Expanded Label	OBO Ontology ID	Markers
Glutamatergic	Glutamatergic neuron	glutamatergic neuron	SV2B, SATB2, CELF2, PHACTR1, CHN1, KHDRBS3, RALYL, LDB2, NELL2, KALRN
GABAergic	GABAergic neuron	GABAergic neuron	GAD1, GRIP1, KAZN, KCNC2, TCF4, GAD2, MAF, SLC6A1, RBMS3, NHS
Non-Neuronal	Non-neuronal cell	animal cell	QKI, ZBTB20, NCKAP5, PTGDS, PHLPP1, CDH20, PPP2R2B, PDE4B, DOCK1, MAP4K4

subclass

Label	Expanded Label	OBO Ontology ID	Markers
Astro	Astrocyte	astrocyte	SLC1A2, ADGRV1, SLC1A3, GPC5, RNF219.AS1, ARHGAP24, CST3, HPSE2, AQP4, COL5A3
Endo	Endothelial cell	endothelial cell	EBF1, ABCG2, CLDN5, FLI1, LEF1, EMCN, IFI27, HLA.E, ADGRL4, CLEC3B
L2/3 IT	Layer 2-3 glutamatergic neuron, intratelencephalon-projecting	glutamatergic neuron	CBLN2, EPHA6, LAMA2, CNTN5, PDZD2, CUX2, RASGRF2, FAM19A1, LINC01378, CA10
L5 ET	Layer 5 glutamatergic neuron, extratelencephalon-projecting	glutamatergic neuron	COL5A2, FAM19A1, VAT1L, COL24A1, CBLN2, NRP1, PTCHD1.AS, NRG1, HOMER1, SLC35F3
L5 IT	Layer 5 glutamatergic neuron, intratelencephalon-projecting	glutamatergic neuron	FSTL4, CNTN5, RORB, FSTL5, IL1RAPL2, CHN2, TOX, CPNE4, CADPS2, POU6F2
L5/6 NP	Layer 5-6 glutamatergic neuron, near-projecting	glutamatergic neuron	TSHZ2, NPSR1.AS1, HTR2C, ITGA8, ZNF385D, ASIC2, CDH6, CRYM, NXPH2, CPNE4
L6 CT	Layer 6 glutamatergic neuron, corticothalamic-projecting	glutamatergic neuron	ADAMTSL1, KIAA1217, SORCS1, HS3ST4, TRPM3, TOX, SEMA3E, EGFEM1P, MEIS2, SEMA5A
L6 IT	Layer 6 glutamatergic neuron, intratelencephalon-projecting	glutamatergic neuron	PTPRK, PDZRN4, CDH9, THEMIS, FSTL5, CDH13, CDH12, CBLN2, LY86.AS1, MLIP
L6 IT Car3	Layer 6 Car3+ glutamatergic neuron, intratelencephalon-projecting	glutamatergic neuron	THEMIS, RNF152, NTNG2, STK32B, KCNMB2, GAS2L3, OLFML2B, POSTN, B3GAT2, NR4A2
L6b	Layer 6b glutamatergic neuron	glutamatergic neuron	HS3ST4, KCNMB2, MDFIC, C10orf11, NTM, CDH9, MARCH1, TLE4, FOXP2, KIAA1217
Lamp5	Lamp5+ GABAergic neuron	GABAergic neuron	FGF13, PTPRT, PRELID2, GRIA4, RELN, PTCHD4, EYA4, MYO16, FBXL7, LAMP5
Micro-PVM	microglia / perivascular macrophage	perivascular macrophage	DOCK8, P2RY12, APBB1IP, FYB, PTPRC, TBXAS1, CX3CR1, BLNK, SLCO2B1, CSF1R
Oligo	oligodendrocyte	oligodendrocyte	PLP1, ST18, CTNNA3, MBP, MOBP, RNF220, NCKAP5, ENPP2, QKI, SLC44A1
OPC	oligodendrocyte precursor cell	oligodendrocyte precursor cell	VCAN, PDGFRA, OLIG1, SMOC1, COL9A1, STK32A, BCAS1, FERMT1, BCHE, ZCCHC24
Pvalb	Pvalb+ GABAergic neuron	GABAergic neuron	ADAMTS17, ERBB4, DPP10, ZNF804A, MYO16, BTBD11, GRIA4, SLIT2, SDK1, PVALB
Sncg	Sncg+ GABAergic neuron	GABAergic neuron	CNR1, SLC8A1, ASIC2, CXCL14, MAML3, ADARB2, NPAS3, CNTN5, FSTL5, SNCG
Sst	Sst+ GABAergic neuron	GABAergic neuron	GRIK1, RALYL, SST, TRHDE, GRID2, NXPH1, COL25A1, SLC8A1, SOX6, ST6GALNAC5
Sst Chodl	Sst+ Chodl+ GABAergic neuron	GABAergic neuron	NPY, FAM46A, STAC, OTOF, NPY2R, CRHBP, ANKRD34B, NOS1, SST, CHODL
Vip	Vip+ GABAergic neuron	GABAergic neuron	GALNTL6, LRP1B, VIP, GRM7, KCNT2, THSD7A, ERBB4, SYNPR, ADARB2, SLC24A3
VLMC	vascular lepotomeningeal cell	brain pericyte	COLEC12, ITIH5, COL1A2, TBX18, EBF1, C7, COL6A2, SRPX2, FLVCR2, FMO2

cluster

Label	Expanded Label	OBO Ontology ID	Markers
Astro L1 FGFR3 SERPINI2	FGFR3+ SERPINI2+ Layer 1 astrocyte	astrocyte	GFAP, PDE4D, ID3, CTNNA2, FGFR3, GRM3, GJA1, TMEM132C, ETNPPL, SERPINI2
Astro L1-6 FGFR3 AQP1	FGFR3+ AQP1+ Layer 1-6 astrocyte	astrocyte	GFAP, TNC, GJA1, ADAMTSL3, RASL12, LINC00609, ANGPTL4, FGFR3, AQP1
Astro L1-6 FGFR3 PLCG1	FGFR3+ PLCG1+ Layer 1-6 astrocyte	astrocyte	FGFR3, HPSE2, GJA1, PLCB1, SLCO1C1, COL5A3, ETNPPL, NDRG2, PRODH, PLCG1
Endo L2-5 NOSTRIN SRGN	NOSTRIN+ SRGN+ Layer 2-5 endothelial cell	endothelial cell	EBF1, CLDN5, ABCG2, FLI1, LEF1, IFI27, EMCN, HLA-E, NOSTRIN, SRGN
Exc L2 LAMP5 KCNG3	LAMP5+ KCNG3+ Layer 2 glutamatergic neuron	glutamatergic neuron	SEMA3C, PARD3, GRM1, SPON1, TMEM196, PALMD, LINGO2, ADRA1A, LAMP5, KCNG3
Exc L2 LINC00507 ATP7B	LINC00507+ ATP7B+ Layer 2 glutamatergic neuron	glutamatergic neuron	OR1J2, KCNK2, ACVR1C, SEMA5A, BDNF, ITGB8, CHRM2, RASAL1, LINC00507, ATP7B
Exc L2 LINC00507 GLRA3	LINC00507+ GLRA3+ Layer 2 glutamatergic neuron	glutamatergic neuron	LRRTM4, MAN1A1, LINC01378, DMD, ENC1, KIAA1217, DGKB, SLIT3, LINC00507, GLRA3
Exc L2-3 LINC00507 DSG3	LINC00507+ DSG3+ Layer 2-3 glutamatergic neuron	glutamatergic neuron	BDNF, BMPR1B, ADCYAP1, SV2C, NOX4, CBLN2, SERPINE2, DLGAP1, LINC00507, DSG3
Exc L2-3 RORB CCDC68	RORB+ CCDC68+ Layer 2-3 glutamatergic neuron	glutamatergic neuron	COL5A2, LRRC4C, RORB, PTPRZ1, NTNG1, LINGO2, VIPR2, CNTNAP5, COBLL1, CCDC68
Exc L2-3 RORB PTPN3	RORB+ PTPN3+ Layer 2-3 glutamatergic neuron	glutamatergic neuron	RORB, GRIA1, COL5A2, PEX5L, RMST, GRM1, NTNG1, GULP1, POU6F2, PTPN3
Exc L2-3 RORB RTKN2	RORB+ RTKN2+ Layer 2-3 glutamatergic neuron	glutamatergic neuron	RMST, GALNT13, COL5A2, ROBO1, ZBTB7C, STXBP6, SCHLAP1, EPHA6, RORB, RTKN2
Exc L3 LAMP5 CARM1P1	LAMP5+ CARM1P1+ Layer 3 glutamatergic neuron	glutamatergic neuron	SV2C, RYR2, SYT2, SLC38A11, BMPR1B, FMN1, MOXD1, PLXDC2, LAMP5, CARM1P1
Exc L3 RORB OTOGL	RORB+ OTOGL+ Layer 3 glutamatergic neuron	glutamatergic neuron	COL22A1, LINCR-0002, PLCH1, RMST, DPP10, TSHZ2, NCAM2, RORB, CPNE4, OTOGL
Exc L3 THEMIS ENPEP	THEMIS+ ENPEP+ Layer 3 glutamatergic neuron	glutamatergic neuron	THEMIS, ITPR2, LAMA2, ASIC2, FAP, KIAA1217, LDB2, PPARGC1A, GAP43, ENPEP
Exc L3-5 FEZF2 ASGR2	FEZF2+ ASGR2+ Layer 3-5 glutamatergic neuron	glutamatergic neuron	OR51E2, SORCS1, LINC00922, LINGO2, ERG, SEMA6D, GFRA1, RASAL2, FEZF2, ASGR2
Exc L3-5 FEZF2 LINC01107	FEZF2+ LINC01107+ Layer 3-5 glutamatergic neuron	glutamatergic neuron	OR51E2, BTNL9, SGCZ, EDNRB, VWC2L, ADGRG6, PCDH11X, LINC00922, FEZF2, LINC01107
Exc L3-5 RORB LAMA4	RORB+ LAMA4+ Layer 3-5 glutamatergic neuron	glutamatergic neuron	TRPC3, DACH1, PLCH1, RORB, CUX2, ANTXR2, IL1RAP, LAMA4, RORA, RMST
Exc L3-5 RORB LINC01202	RORB+ LINC01202+ Layer 3-5 glutamatergic neuron	glutamatergic neuron	LINCR-0002, PCDH7, TSHZ2, HS3ST4, RORB, IL1RAPL2, TPBG, HTR1F, EPHA3, LINC01202
Exc L3-5 RORB LNX2	RORB+ LNX2+ Layer 3-5 glutamatergic neuron	glutamatergic neuron	CADPS2, FAM19A1, LRRK1, NTM, PCP4, RGS6, COL11A1, GRM7, CHN2, LNX2
Exc L3-5 RORB RPRM	RORB+ RPRM+ Layer 3-5 glutamatergic neuron	glutamatergic neuron	MKX, RPRM, RXFP1, CCK, RORB, SLC9A9, FSTL4, SPARCL1, IPCEF1, SEMA3E
Exc L3-5 RORB TNNT2	RORB+ TNNT2+ Layer 3-5 glutamatergic neuron	glutamatergic neuron	LINCR-0002, IL1RAPL2, THEMIS, CPNE4, FSHR, LAMA2, UTRN, RORB, TSHZ2, TNNT2
Exc L5 FEZF2 CSN1S1	FEZF2+ CSN1S1+ Layer 5 glutamatergic neuron	glutamatergic neuron	ANGPT1, KCNIP1, SULF2, KCNG2, ANKRD26P3, NETO1, PAPPA, GRM1, FEZF2, CSN1S1
Exc L5 FEZF2 NREP-AS1	FEZF2+ NREP-AS1+ Layer 5 glutamatergic neuron	glutamatergic neuron	CD36, NWD2, ADAMTS12, GPC5, LINC01507, GRIK1, FER1L6-AS2, MAST4, FEZF2, NREP-AS1
Exc L5 FEZF2 PKD2L1	FEZF2+ PKD2L1+ Layer 5 glutamatergic neuron	glutamatergic neuron	FER1L6-AS2, GPC5, NPSR1-AS1, PDE1A, ROBO3, TRPM3, NPSR1, RALYL, FEZF2, PKD2L1
Exc L5 FEZF2 RNF144A-AS1	FEZF2+ RNF144A-AS1+ Layer 5 glutamatergic neuron	glutamatergic neuron	IFNG-AS1, GPC6, CD36, GRIK1, GDNF-AS1, PVT1, NPSR1, LRRC4C, FEZF2, RNF144A-AS1
Exc L5 RORB MED8	RORB+ MED8+ Layer 5 glutamatergic neuron	glutamatergic neuron	NPFFR2, COL22A1, FAM19A2, IL1RAPL2, FNBP1L, RXFP1, LRRC7, RORB, KCNH5, MED8
Exc L5 THEMIS FGF10	THEMIS+ FGF10+ Layer 5 glutamatergic neuron	glutamatergic neuron	NPFFR2, TESPA1, CRH, RCAN2, THEMIS, NELL1, BDNF, CA10, RXFP1, FGF10
Exc L5 THEMIS LINC01116	THEMIS+ LINC01116+ Layer 5 glutamatergic neuron	glutamatergic neuron	LINC01060, HCRTR2, SEMA3E, DPP4, THEMIS, LINC01116
Exc L5 THEMIS RGPD6	THEMIS+ RGPD6+ Layer 5 glutamatergic neuron	glutamatergic neuron	SEMA3E, TRPC6, SEMA5A, THEMIS, DDR2, RNF152, LINC01060, EGFEM1P, RGPD6
Exc L5 THEMIS SLC22A18	THEMIS+ SLC22A18+ Layer 5 glutamatergic neuron	glutamatergic neuron	KCNG2, GRIN3A, CCBE1, COL21A1, THEMIS, EPHA6, MKX, LDLRAD4, HS3ST2, SLC22A18
Exc L5 THEMIS VILL	THEMIS+ VILL+ Layer 5 glutamatergic neuron	glutamatergic neuron	TNC, SAMD5, NPFFR2, NELL1, FAP, LRP1B, DACH1, CDH6, THEMIS, VILL
Exc L5-6 FEZF2 C9orf135-AS1	FEZF2+ C9orf135-AS1+ Layer 5-6 glutamatergic neuron	glutamatergic neuron	SEMA3E, OPCML, EGFEM1P, LRRTM4, SEMA5A, RALYL, HS3ST4, PRKD1, FEZF2, C9ORF135-AS1
Exc L5-6 FEZF2 CFTR	FEZF2+ CFTR+ Layer 5-6 glutamatergic neuron	glutamatergic neuron	NR4A2, TMEM244, CTGF, CALCRL, MCTP2, CLDN1, ERG, EPB41L4A, FEZF2, CFTR
Exc L5-6 FEZF2 FILIP1L	FEZF2+ FILIP1L+ Layer 5-6 glutamatergic neuron	glutamatergic neuron	CALCRL, ALDOA, SEMA3E, XIST, SEMA5A, EGFEM1P, FEZF2, FILIP1L
Exc L5-6 FEZF2 IFNG-AS1	FEZF2+ IFNG-AS1+ Layer 5-6 glutamatergic neuron	glutamatergic neuron	CD36, NRXN3, IFNG-AS1, KCNQ5, FER1L6-AS2, PCDH11X, HTR2C, DGKI, NPSR1-AS1, FEZF2
Exc L5-6 FEZF2 LPO	FEZF2+ LPO+ Layer 5-6 glutamatergic neuron	glutamatergic neuron	GADL1, ALCAM, MYHAS, PCED1B, CALCRL, LINC01105, FER1L6-AS2, ZNF385D-AS2, FEZF2, LPO
Exc L5-6 FEZF2 OR1L8	FEZF2+ OR1L8+ Layer 5-6 glutamatergic neuron	glutamatergic neuron	SYT6, KCNC2, MCTP2, CTNND2, SULF1, KIAA1217, RBM20, PLXDC2, FEZF2, OR1L8
Exc L5-6 FEZF2 SH2D1B	FEZF2+ SH2D1B+ Layer 5-6 glutamatergic neuron	glutamatergic neuron	SEMA3E, PRKY, DDR2, USP9Y, DCHS2, NLGN4Y, SEMA5A, DDX3Y, FEZF2, SH2D1B
Exc L5-6 THEMIS SMYD1	THEMIS+ SMYD1+ Layer 5-6 glutamatergic neuron	glutamatergic neuron	NR4A2, KCNMB2, GAS2L3, SEMA6D, ABCC3, TSHZ2, NTNG2, EDIL3, THEMIS, SMYD1
Exc L5-6 THEMIS TNFAIP6	THEMIS+ TNFAIP6+ Layer 5-6 glutamatergic neuron	glutamatergic neuron	THEMIS, CDH9, HS3ST2, CDH12, MKX, SPOCK1, TSHZ2, KCTD16, RXFP1, TNFAIP6
Exc L6 FEZF2 FFAR4	FEZF2+ FFAR4+ Layer 6 glutamatergic neuron	glutamatergic neuron	SLC15A5, OLFML2B, CTGF, FER1L6-AS2, FEZF2, FFAR4
Exc L6 FEZF2 KLK7	FEZF2+ KLK7+ Layer 6 glutamatergic neuron	glutamatergic neuron	MDFIC, ASIC2, ZFHX3, CHN2, SEMA3D, SDK1, MCTP2, COL24A1, FEZF2, KLK7
Exc L6 FEZF2 PDYN	FEZF2+ PDYN+ Layer 6 glutamatergic neuron	glutamatergic neuron	SLC15A5, ADGRF2, PDYN, CABP7, SYT6, SEMA3D, ERG, FEZF2
Exc L6 FEZF2 POGK	FEZF2+ POGK+ Layer 6 glutamatergic neuron	glutamatergic neuron	CTGF, CHRM2, CDC14A, MGAT4C, SEMA3D, PLCXD3, NPFFR2, KCNK2, FEZF2, POGK
Exc L6 FEZF2 PROKR2	FEZF2+ PROKR2+ Layer 6 glutamatergic neuron	glutamatergic neuron	SLC15A5, CTGF, VCAN, ADD3-AS1, FEZF2, PROKR2
Exc L6 THEMIS LINC00343	THEMIS+ LINC00343+ Layer 6 glutamatergic neuron	glutamatergic neuron	THEMIS, ROBO1, ADAMTS3, GULP1, ANKRD30B, TRPM3, DCHS2, SEMA3A, SEC24D, LINC00343
Exc L6 THEMIS SLN	THEMIS+ SLN+ Layer 6 glutamatergic neuron	glutamatergic neuron	THEMIS, PDE7B, LINCR-0002, LINC01378, GBE1, GRIA4, ARHGEF28, C8orf34, FRMD6, SLN
Exc L6 THEMIS SNTG2	THEMIS+ SNTG2+ Layer 6 glutamatergic neuron	glutamatergic neuron	THEMIS, THSD7B, EDIL3, SNTG2, TENM4, ADAMTS3, CLSTN2, LY86-AS1, CNTN5, DCHS2
Inh L1 LAMP5 BMP2	LAMP5+ BMP2+ Layer 1 GABAergic neuron	GABAergic neuron	NDNF, SYT1, NPNT, ALK, CXCL14, RBFOX1, RELN, PBX1, LAMP5, BMP2
Inh L1 LAMP5 NMBR	LAMP5+ NMBR+ Layer 1 GABAergic neuron	GABAergic neuron	NDNF, MAGI2, PCDH18, PDE4B, NMBR, PRELID2, NPNT, EPHA6, CXCL14, LAMP5
Inh L1 LAMP5 PVRL2	LAMP5+ PVRL2+ Layer 1 GABAergic neuron	GABAergic neuron	NDNF, UNC13C, MYH11, TMEM132D, KIT, PCDH9, RELN, CLSTN2, LAMP5, PVRL2
Inh L1 LAMP5 RAB11FIP1	LAMP5+ RAB11FIP1+ Layer 1 GABAergic neuron	GABAergic neuron	MYH11, CDH9, KIT, RELN, ADAMTS20, CACNA2D1, KIRREL, CDH11, LAMP5, RAB11FIP1
Inh L1 PAX6 CHRFAM7A	PAX6+ CHRFAM7A+ Layer 1 GABAergic neuron	GABAergic neuron	LINC01539, FSTL5, NDNF, CNTNAP4, CXCL14, ZMAT4, NR2F2-AS1, LIMCH1, PAX6, CHRFAM7A
Inh L1 PAX6 MIR101-1	PAX6+ MIR101-1+ Layer 1 GABAergic neuron	GABAergic neuron	TGFBR2, SPHKAP, SP8, ZNF804A, CDH23, PDZRN4, GPC3, ALK, PAX6, MIR101-1
Inh L1 PVALB SST ASIC4	PVALB+ SST ASIC4+ Layer 1 GABAergic neuron	GABAergic neuron	ADAM33, RXRG, NDNF, EFEMP1, PVALB, SST ASIC4
Inh L1 SST DEFB108B	SST+ DEFB108B+ Layer 1 GABAergic neuron	GABAergic neuron	PCDH18, ARPP21, NR2F2, THSD7A, SLC10A4, SYNPR, SYT10, CPE, SST, DEFB108B
Inh L1 SST P4HA3	SST+ P4HA3+ Layer 1 GABAergic neuron	GABAergic neuron	CALCB, RXRG, NDNF, ADAM33, SST, P4HA3
Inh L1 VIP KLHDC8B	VIP+ KLHDC8B+ Layer 1 GABAergic neuron	GABAergic neuron	VIP, GPC6, CHRNA2, ST6GALNAC5, NDNF, ADAM12, TRPC7, CNTN5, PROX1-AS1, KLHDC8B
Inh L1-2 PVALB CDK20	PVALB+ CDK20+ Layer 1-2 GABAergic neuron	GABAergic neuron	TAC1, SYT10, SYTL5, NRP2, NDST4, KCNT2, MYO5B, FGD5, PVALB, CDK20
Inh L1-2 SST CCNJL	SST+ CCNJL+ Layer 1-2 GABAergic neuron	GABAergic neuron	CALB1, ALK, DKK2, ALCAM, CMTM8, ERBB4, MYO5B, TMEM132C, SST, CCNJL
Inh L1-2 SST CLIC6	SST+ CLIC6+ Layer 1-2 GABAergic neuron	GABAergic neuron	TAC1, NDST4, NKAIN2, SYTL5, SGCZ, CMTM8, NEDD4, FBN2, SST, CLIC6
Inh L1-2 SST PRRT4	SST+ PRRT4+ Layer 1-2 GABAergic neuron	GABAergic neuron	PENK, PDE4B, CALB1, ARHGAP10, HPGD, ITPR2, SST, ZNF385B, STK32B, PRRT4
Inh L1-2 VIP EXPH5	VIP+ EXPH5+ Layer 1-2 GABAergic neuron	GABAergic neuron	SHISA8, LINGO2, VIP, BTBD11, SLC22A3, ASIC2, FLT1, FAM19A2, ASIC4, EXPH5
Inh L1-2 VIP HTR3A	VIP+ HTR3A+ Layer 1-2 GABAergic neuron	GABAergic neuron	HTR3A, RXFP1, VWA5B1, CSMD1, NR2F2-AS1, EDIL3, EDNRA, INPP4B, COL4A4, VIP
Inh L1-2 VIP PTGER3	VIP+ PTGER3+ Layer 1-2 GABAergic neuron	GABAergic neuron	NPR3, PTGER3, NR2F2, EDNRA, PROX1-AS1, VIP
Inh L1-2 VIP SCML4	VIP+ SCML4+ Layer 1-2 GABAergic neuron	GABAergic neuron	SCML4, ADAMTS19, COL9A1, NELL1, CALB2, SLC6A11, NR2F2, KCNH8, NR2F2-AS1, VIP
Inh L1-2 VIP WNT4	VIP+ WNT4+ Layer 1-2 GABAergic neuron	GABAergic neuron	NR2F2, CCK, CFAP47, CNR1, GABRG1, SORCS3, NECAB2, ROBO1, VIP, WNT4
Inh L1-3 SST FAM20A	SST+ FAM20A+ Layer 1-3 GABAergic neuron	GABAergic neuron	CCDC109B, LINGO2, SST, CDH12, SYTL5, NCAM2, SOX6, TAC1, KIRREL3, FAM20A
Inh L1-3 VIP CBLN1	VIP+ CBLN1+ Layer 1-3 GABAergic neuron	GABAergic neuron	SLC22A3, SOX2-OT, CBLN1, LINC01322, TAC3, ZBTB20, SHISA8, SOX5, HTR2C, VIP
Inh L1-3 VIP CHRNA2	VIP+ CHRNA2+ Layer 1-3 GABAergic neuron	GABAergic neuron	NPR3, DPP6, CHRNA2, CDH20, PPP1R1C, CNTNAP4, CALB2, SLC7A11, VIP, LYPD1
Inh L1-3 VIP FNDC1	VIP+ FNDC1+ Layer 1-3 GABAergic neuron	GABAergic neuron	ADAM33, EPHA7, SHISA8, ASIC4, PRR16, LINC01170, ADAMTS19, NPR3, VIP, FNDC1
Inh L1-3 VIP HSPB6	VIP+ HSPB6+ Layer 1-3 GABAergic neuron	GABAergic neuron	SLC22A3, KCNQ5, VIP, CNTN5, SHISA8, SEMA6D, TAC3, NPAS3, PTHLH, HSPB6
Inh L1-5 VIP CD27-AS1	VIP+ CD27-AS1+ Layer 1-5 GABAergic neuron	GABAergic neuron	CALB2, ZDHHC2, IQGAP2, TOX, VIP, KLHL1, NR2E1, FAT3, SMOC1, CD27-AS1
Inh L1-5 VIP LINC01013	VIP+ LINC01013+ Layer 1-5 GABAergic neuron	GABAergic neuron	PTGFR, INPP4B, PCDH18, DAB1, NR2F2, PRELID2, NR2F2-AS1, VIP, LINC01013
Inh L1-5 VIP PHLDB3	VIP+ PHLDB3+ Layer 1-5 GABAergic neuron	GABAergic neuron	TAC3, ASIC4, SLC22A3, CHRNA2, VIP, PHLDB3
Inh L1-5 VIP SMOC1	VIP+ SMOC1+ Layer 1-5 GABAergic neuron	GABAergic neuron	VIP, CNR1, CHRNA2, GULP1, TAC3, LRP1B, SMOC1, SHISA9, SLC22A3, NPR3
Inh L1-6 LAMP5 AARD	LAMP5+ AARD+ Layer 1-6 GABAergic neuron	GABAergic neuron	KIT, CDH13, PKP2, PIP5K1B, EYA4, PRELID2, ROR2, LRP1B, LAMP5, AARD
Inh L1-6 LAMP5 CA1	LAMP5+ CA1+ Layer 1-6 GABAergic neuron	GABAergic neuron	SFRP1, TAC1, KIRREL, OPCML, CA8, PRKG1, NOS1, HS3ST4, LAMP5, CA1
Inh L1-6 LAMP5 NES	LAMP5+ NES+ Layer 1-6 GABAergic neuron	GABAergic neuron	ROR2, KCNMB2, KIT, CCBE1, EYA4, MEIS2, PDGFRA, C8orf34, LAMP5, NES
Inh L1-6 PVALB COL15A1	PVALB+ COL15A1+ Layer 1-6 GABAergic neuron	GABAergic neuron	NPNT, HTR1F, COL15A1, GULP1, PTHLH, FSTL5, ADAMTS5, CNTNAP4, FAM19A4, PVALB
Inh L1-6 SST NPY	SST+ NPY+ Layer 1-6 GABAergic neuron	GABAergic neuron	NPY, FAM46A, STAC, OTOF, ANKRD34B, CRHBP, NPY2R, NOS1, CCDC109B, SST
Inh L1-6 VIP SLC7A6OS	VIP+ SLC7A6OS+ Layer 1-6 GABAergic neuron	GABAergic neuron	DYNAP, RASGRF2, FREM2, LINC01539, PBX3, TP53I11, PCP4, VIP, PDLIM5, SLC7A6OS
Inh L2 PAX6 FREM2	PAX6+ FREM2+ Layer 2 GABAergic neuron	GABAergic neuron	SLC17A8, PDE3A, DOCK8, GRIK1, IGFBP7, PCLO, FREM2, YJEFN3, NR1H4, PAX6
Inh L2 PVALB FRZB	PVALB+ FRZB+ Layer 2 GABAergic neuron	GABAergic neuron	CRHBP, ARHGAP42, KAL1, MYO5B, PVALB, FRZB
Inh L2 VIP SLC6A16	VIP+ SLC6A16+ Layer 2 GABAergic neuron	GABAergic neuron	NPR3, CDH8, NR2F2, CALB2, NR2F2-AS1, VIP, SLC6A16
Inh L2-3 SST NMU	SST+ NMU+ Layer 2-3 GABAergic neuron	GABAergic neuron	MYO5B, ERBB4, SOX6, FAM19A2, PIEZO2, MYO16, NOS1, NXPH1, SST, NMU
Inh L2-5 PVALB HHIPL1	PVALB+ HHIPL1+ Layer 2-5 GABAergic neuron	GABAergic neuron	MEPE, EPHA3, PVALB, POU6F2, POSTN, GPR149, PLEKHG1, TAC1, ADAMTS20, HHIPL1
Inh L2-5 PVALB RPH3AL	PVALB+ RPH3AL+ Layer 2-5 GABAergic neuron	GABAergic neuron	BTBD11, LRP1B, ADAMTS17, MSRA, MYO5B, BCL11A, RNF144B, LINC00403, PVALB, RPH3AL
Inh L2-5 VIP BSPRY	VIP+ BSPRY+ Layer 2-5 GABAergic neuron	GABAergic neuron	CHRNA2, ST18, VIP, SFMBT2, SHISA8, CA10, RGS16, PCDH7, SLC22A3, BSPRY
Inh L2-5 VIP SOX11	VIP+ SOX11+ Layer 2-5 GABAergic neuron	GABAergic neuron	PENK, PCDH18, PTGFR, CALB2, VIP, SOX11
Inh L3 PVALB SAMD13	PVALB+ SAMD13+ Layer 3 GABAergic neuron	GABAergic neuron	PVALB, TOX, SULF1, EPHA3, TAC1, HOMER3, POSTN, PHACTR2, RNF144B, SAMD13
Inh L3-5 PVALB ISG20	PVALB+ ISG20+ Layer 3-5 GABAergic neuron	GABAergic neuron	PVALB, PHACTR2, LCNL1, EPHA4, POSTN, GRM8, SULF1, RBMS3, RNF144B, ISG20
Inh L3-5 SST CDH3	SST+ CDH3+ Layer 3-5 GABAergic neuron	GABAergic neuron	LINC01324, IGFBP5, OTOF, POSTN, QRFPR, LINC01399, EDNRA, GDA, SST, CDH3
Inh L3-5 SST GGTLC3	SST+ GGTLC3+ Layer 3-5 GABAergic neuron	GABAergic neuron	ADGRG6, ZNF385D, SST, SOX5, EPSTI1, MGAT4C, NMU, ROBO1, GRIK1, GGTLC3
Inh L3-5 SST OR5AH1P	SST+ OR5AH1P+ Layer 3-5 GABAergic neuron	GABAergic neuron	NMU, EPHA6, SST, ADAMTS9-AS2, EDNRA, PCDH9, PAWR, PCSK5, ADAMTS5, OR5AH1P
Inh L3-5 VIP HS3ST3A1	VIP+ HS3ST3A1+ Layer 3-5 GABAergic neuron	GABAergic neuron	SLC22A3, GRM8, IQGAP2, THSD7B, PTHLH, UNC13C, CHRNA2, CDH13, VIP, HS3ST3A1
Inh L3-5 VIP IGDCC3	VIP+ IGDCC3+ Layer 3-5 GABAergic neuron	GABAergic neuron	EREG, NECAB1, TAC3, ANKS1B, SLC22A3, GRIA1, VIP, PTPRD, IQGAP2, IGDCC3
Inh L3-5 VIP TAC3	VIP+ TAC3+ Layer 3-5 GABAergic neuron	GABAergic neuron	TAC3, NBEA, PPP1R1C, ARHGAP18, VIP, EGFEM1P, SLC22A3, NCAM2, IQGAP2, NR2F2-AS1
Inh L3-6 PAX6 LINC01497	PAX6+ LINC01497+ Layer 3-6 GABAergic neuron	GABAergic neuron	SCGN, EGFEM1P, PALMD, TMEM132D, EYA1, C8orf34, TNS3, KCNIP4, PAX6, LINC01497
Inh L3-6 VIP UG0898H09	VIP+ UG0898H09+ Layer 3-6 GABAergic neuron	GABAergic neuron	PCDH18, VIP, CHRNA2, PTGFR, LINC01411, UG0898H09
Inh L3-6 VIP ZIM2-AS1	VIP+ ZIM2-AS1+ Layer 3-6 GABAergic neuron	GABAergic neuron	PENK, PTGFR, VGLL3, ROR2, VIP, ZIM2-AS1
Inh L5 PVALB LRIG3	PVALB+ LRIG3+ Layer 5 GABAergic neuron	GABAergic neuron	HGF, TMTC2, PVALB, ROBO1, GPR149, RIT2, PLCE1, LHFPL3, MEPE, LRIG3
Inh L5 SST RPL35AP11	SST+ RPL35AP11+ Layer 5 GABAergic neuron	GABAergic neuron	SST, ADCY2, EDNRA, GRIK2, QRFPR, DMD, NMU, GRIK1, PAWR, RPL35AP11
Inh L5-6 LAMP5 CRABP1	LAMP5+ CRABP1+ Layer 5-6 GABAergic neuron	GABAergic neuron	SFRP1, COL25A1, FREM2, PRELID2, PDGFRA, KCNIP4, KIRREL, FBN2, LAMP5, CRABP1
Inh L5-6 PVALB FAM150B	PVALB+ FAM150B+ Layer 5-6 GABAergic neuron	GABAergic neuron	FAM150B, GPC5, PALMD, SAMD3, MYBPC1, PDZRN4, ANKRD34B, TMEM200A, GPR149, PVALB
Inh L5-6 PVALB GAPDHP60	PVALB+ GAPDHP60+ Layer 5-6 GABAergic neuron	GABAergic neuron	WNT16, CDH9, MYBPC1, NCAM2, BCHE, KIRREL3, WIF1, PVALB, GAPDHP60
Inh L5-6 PVALB KCNIP2	PVALB+ KCNIP2+ Layer 5-6 GABAergic neuron	GABAergic neuron	NPFFR2, GULP1, THSD7B, CDH8, WIF1, SORCS3, LYPD6, PAWR, PVALB, KCNIP2
Inh L5-6 PVALB MEPE	PVALB+ MEPE+ Layer 5-6 GABAergic neuron	GABAergic neuron	WNT16, PCDH9, MEPE, MCTP1, ANKRD34B, PRKG1, MYBPC1, GALNTL6, CPED1, PVALB
Inh L5-6 PVALB SST CRHR2	PVALB+ SST CRHR2+ Layer 5-6 GABAergic neuron	GABAergic neuron	HPGD, ADAMTS17, FAM89A, CNTNAP4, FLT1, DGKB, TAC1, GHR, PVALB, SST CRHR2
Inh L5-6 PVALB ZFPM2-AS1	PVALB+ ZFPM2-AS1+ Layer 5-6 GABAergic neuron	GABAergic neuron	ZFPM2-AS1, WIF1, TMTC2, POSTN, SEMA6D, LYPD6, GRIK2, SULF1, PCDH15, PVALB
Inh L5-6 SST BEAN1	SST+ BEAN1+ Layer 5-6 GABAergic neuron	GABAergic neuron	FBN2, PCDH11X, THSD7B, CDH10, KLHL14, CDH9, SST, DPP10, HGF, BEAN1
Inh L5-6 SST C4orf26	SST+ C4orf26+ Layer 5-6 GABAergic neuron	GABAergic neuron	ADAMTS20, LINGO2, FBN2, PLXDC2, KLHL14, PDE1C, DCN, PCDH17, HGF, C4ORF26
Inh L5-6 SST DNAJC14	SST+ DNAJC14+ Layer 5-6 GABAergic neuron	GABAergic neuron	HGF, PLCE1, FBN2, PTPRK, ITGA1, TRPC5, GPR149, LRRC4C, SST, DNAJC14
Inh L5-6 SST FBN2	SST+ FBN2+ Layer 5-6 GABAergic neuron	GABAergic neuron	FBN2, ZNF385D, FAM129A, CSMD1, KLHL14, EFNA5, ADAMTS20, MGAT4C, THSD7B, SST
Inh L5-6 SST ISX	SST+ ISX+ Layer 5-6 GABAergic neuron	GABAergic neuron	SST, CSGALNACT1, NR2F2-AS1, CNTNAP4, DKK2, TRPM3, PREX2, GRM7, SOX6, ISX
Inh L5-6 SST KLHL1	SST+ KLHL1+ Layer 5-6 GABAergic neuron	GABAergic neuron	SLC17A8, ADGRL2, DCN, ROBO1, KLHL14, SOX6, ADAMTS20, NDST4, UNC13C, KLHL1
Inh L5-6 SST PAWR	SST+ PAWR+ Layer 5-6 GABAergic neuron	GABAergic neuron	NDST4, LRRTM4, NMU, PTPRT, DCN, ALCAM, KLHL14, GALNTL6, TMCC3, SST
Inh L5-6 SST PIK3CD	SST+ PIK3CD+ Layer 5-6 GABAergic neuron	GABAergic neuron	ADGRG6, NLGN1, KLHL14, LRFN5, NMU, ARHGAP10, NDST4, NEGR1, SST, PIK3CD
Inh L5-6 VIP COL4A3	VIP+ COL4A3+ Layer 5-6 GABAergic neuron	GABAergic neuron	VIP, C11orf1, SLC22A3, GRM8, ABI3BP, CBLN4, NPR3, UNC13C, VCAN, COL4A3
Inh L6 SST TH	SST+ TH+ Layer 6 GABAergic neuron	GABAergic neuron	KIAA1024, ERBB4, RAB3B, GPC5, SST, PTPRK, GABRG1, KLHL29, WIF1, TH
Micro L1-6 TYROBP CD74	TYROBP+ CD74+ Layer 1-6 microglia	microglial cell	DOCK8, P2RY12, APBB1IP, FYB, TBXAS1, SLCO2B1, CX3CR1, PTPRC, TYROBP, CD74
Oligo L2-6 OPALIN FTH1P3	OPALIN+ FTH1P3+ Layer 2-6 oligodendrocyte	oligodendrocyte	PLP1, ANK3, ST18, NLGN1, RNF220, PLXDC2, ENPP2, OPALIN, FRMD4B, FTH1P3
Oligo L2-6 OPALIN MAP6D1	OPALIN+ MAP6D1+ Layer 2-6 oligodendrocyte	oligodendrocyte	OPALIN, FOLH1, ABCA8, MOG, UGT8, SEPP1, LPAR1, ST18, SLCO1A2, MAP6D1
Oligo L3-6 OPALIN ENPP6	OPALIN+ ENPP6+ Layer 3-6 oligodendrocyte	oligodendrocyte	S100B, RBFOX1, CARNS1, RASGRF1, COL4A5, COL18A1, MOG, USH1C, OPALIN, ENPP6
Oligo L3-6 OPALIN LRP4-AS1	OPALIN+ LRP4-AS1+ Layer 3-6 oligodendrocyte	oligodendrocyte	LINC00982, FGFR3, NKAIN3, OPALIN, SLC1A2, TRIM59, RYR3, GJA1, RNF219-AS1, LRP4-AS1
Oligo L5-6 OPALIN LDLRAP1	OPALIN+ LDLRAP1+ Layer 5-6 oligodendrocyte	oligodendrocyte	PLP1, ANK3, ST18, NLGN1, RNF220, PLXDC2, ENPP2, OPALIN, FRMD4B, LDLRAP1
OPC L1-6 PDGFRA COL20A1	PDGFRA+ COL20A1+ Layer 1-6 oligodendrocyte precursor cell	oligodendrocyte precursor cell	OLIG1, COL9A1, OLIG2, SNX22, NKAIN4, KLRC4-KLRK1, VCAN, PDGFRA, COL20A1, GPR37L1
VLMC L1-5 PDGFRA COLEC12	PDGFRA+ COLEC12+ Layer 1-5 vascular lepotomeningeal cell	brain pericyte	COLEC12, COL1A2, TBX18, ITIH5, COL6A2, SRPX2, C7, FMO2, MFAP4, PDGFRA

cross-species cluster

Label	Expanded Label	OBO Ontology ID	Markers
Astro_1	Astrocyte_1	astrocyte	GFAP, FAM189A2, FGFR3, GJA1, SLC14A1, AGT, ID3, RASL12, AQP4, ANGPTL4
Astro_2	Astrocyte_2	astrocyte	FGFR3, GJA1, SLCO1C1, LINC00982, SDC4, NFATC4, GJB6, PRODH, APOE, SLC1A3
Chandelier	Chandelier neuron	GABAergic neuron	NPNT, COL15A1, PLEKHH2, NOG, PTHLH, ADAMTS5, COL21A1, FAM19A4, HTR1F, SRGAP1
Endo	Endothelial cell	endothelial cell	EBF1, CLDN5, ABCG2, FLI1, LEF1, IFI27, EMCN, HLA-E, ID3, XAF1
L2/3 IT	Layer 2-3 glutamatergic neuron, intratelencephalon-projecting	glutamatergic neuron	PDZD2, CBLN2, MEIS2, CUX2, KIAA1217, ENC1, DAB1, PVRL3, GNAL, LRRTM4
L5 ET_1	Layer 5 glutamatergic neuron, extratelencephalon-projecting_1	glutamatergic neuron	OR51E2, SORCS1, LINC00922, SEMA6D, FAM84B, TMEM2, LINC01036, ST6GALNAC3, VIM-AS1, PQLC2L
L5 ET_2	Layer 5 glutamatergic neuron, extratelencephalon-projecting_2	glutamatergic neuron	BTNL9, EDNRB, SGCZ, LINC00943, VWC2L, HBE1, PCDH11X, TRAPPC3L, CDH8, LINC01069
L5 IT_1	Layer 5 glutamatergic neuron, intratelencephalon-projecting_1	glutamatergic neuron	RORB, SEMA6D, POU6F2, TMEM196, PLCH1, ALCAM, RMST, CUX1, DCC, TRHDE
L5 IT_2	Layer 5 glutamatergic neuron, intratelencephalon-projecting_2	glutamatergic neuron	CHN2, LRRK1, CAMK2D, FSTL4, PDE1C, DNAH6, PCDH7, PRUNE2, ARSJ, TMSB10
L5 IT_3	Layer 5 glutamatergic neuron, intratelencephalon-projecting_3	glutamatergic neuron	HS3ST3B1, RORB, HES1, CSGALNACT1, CRH, PDZRN3, DRD5, CCDC85A, ARHGAP15, TNNT2
L5/6 NP	Layer 5-6 glutamatergic neuron, near-projecting	glutamatergic neuron	NPSR1-AS1, CBLN2, FER1L6-AS2, LUZP2, TLL1, KIAA1217, CALCRL, TLE4, HTR2C, PDE9A
L6 CT_1	Layer 6 glutamatergic neuron, corticothalamic-projecting_1	glutamatergic neuron	SYT6, FAM189A1, VWA2, SPON1, KL, ASTN2, SEMA5A, WBSCR17, ABO, LGR6
L6 CT_2	Layer 6 glutamatergic neuron, corticothalamic-projecting_2	glutamatergic neuron	VWA2, ERBB4, ABO, PRKD1, SH2D1B, ZNF804B, IL4R, ROBO2, DCHS2, ADAMTSL1
L6 IT_1	Layer 6 glutamatergic neuron, intratelencephalon-projecting_1	glutamatergic neuron	LINC00343, TENM3, WISP1, CDH13, ANKRD30B, MEIS2, FRMD6, RASGRF2, TTC6, SEC24D
L6 IT_2	Layer 6 glutamatergic neuron, intratelencephalon-projecting_2	glutamatergic neuron	SH2D1B, EDIL3, TMEM244, CNTN5, ADAMTS16, PDE1A, MYO1D, TSHZ2, ANO3, PXDN
L6 IT_3	Layer 6 glutamatergic neuron, intratelencephalon-projecting_3	glutamatergic neuron	SMYD1, KCNMB2, OLFML2B, RGS12, NR4A2, SLC24A4, LINC00331, RNF152, GAS2L3, PRLR
L6b	Layer 6b glutamatergic neuron	glutamatergic neuron	ZFHX3, C10orf11, MDFIC, MARCH1, SEMA3D, ARSJ, GRIA4, MCTP2, KCNMB2, CTGF
Lamp5_1	Lamp5+ GABAergic neuron_1	GABAergic neuron	NDNF, UNC13C, CPLX3, PCDH9, MYH11, CLSTN2, LINC00298, TMEM132D, FAT1, LAMP5
Lamp5_2	Lamp5+ GABAergic neuron_2	GABAergic neuron	KIT, NTNG1, FBXL7, TOX, NXPH2, NCAM2, PKP2, PTPRG, HAPLN1, LAMP5
Lamp5_3	Lamp5+ GABAergic neuron_3	GABAergic neuron	PDGFD, ITGA8, EYA4, CHST9, SFRP1, SORCS1, KIRREL, UNC5D, PDGFRA, LAMP5
Lamp5_4	Lamp5+ GABAergic neuron_4	GABAergic neuron	SP8, SPHKAP, CDH23, MYO16, NTF3, NXPH1, TGFBR2, ALK, DKFZp686K1684, LAMP5
Lamp5_5	Lamp5+ GABAergic neuron_5	GABAergic neuron	NDNF, COL5A2, CXCL14, HS6ST3, NMBR, ALCAM, BMP6, EPHA6, FAT1, LAMP5
Microglia/PVM	microglia / perivascular macrophage	perivascular macrophage	DOCK8, P2RY12, APBB1IP, FYB, TBXAS1, SLCO2B1, CX3CR1, PTPRC, BLNK, CSF1R
Oligo_1	oligodendrocyte_1	oligodendrocyte	S100B, RBFOX1, COL4A5, RASGRF1, CARNS1, COL18A1, MOG, USH1C, ABCA8, FOLH1
Oligo_2	oligodendrocyte_2	oligodendrocyte	PLP1, LINC00609, ENPP2, PLXDC2, UGT8, OPALIN, BCAS1, NLGN1, CERCAM, DOCK5
OPC	oligodendrocyte precursor cell	oligodendrocyte precursor cell	OLIG1, COL9A1, OLIG2, SNX22, NKAIN4, KLRC4-KLRK1, GPR17, VCAN, COL20A1, GPR37L1
Pvalb_1	Pvalb+ GABAergic neuron_1	GABAergic neuron	ADAMTS17, GULP1, BTBD11, DPP10, RNF144B, THSD7B, CNTNAP3, TENM4, TMEM132C, PVALB
Pvalb_2	Pvalb+ GABAergic neuron_2	GABAergic neuron	WNT16, SHISA8, MYBPC1, PALMD, FAM150B, TRPC6, GPR149, ANKRD34B, GABRG1, PVALB
Sncg_1	Sncg+ GABAergic neuron_1	GABAergic neuron	LINC01539, SORCS1, DYNAP, DSCAM, SP8, PROX1, CHRFAM7A, PDE7B, HIF3A, SNCG
Sncg_2	Sncg+ GABAergic neuron_2	GABAergic neuron	SLC17A8, YJEFN3, DOCK8, ARHGAP6, IGFBP7, RYR2, FREM2, PCLO, CDH23, SNCG
Sncg_3	Sncg+ GABAergic neuron_3	GABAergic neuron	HTR3A, GRM1, VWA5B1, SYNPR, HGFAC, HDAC9, SLITRK6, PTPRM, PLSCR5, SNCG
Sncg_4	Sncg+ GABAergic neuron_4	GABAergic neuron	RXRG, ARPP21, NDNF, SYNPR, NR2F2, FAM19A1, GNG12, THSD7A, ADAM33, SNCG
Sst Chodl	Sst+ Chodl+ GABAergic neuron	GABAergic neuron	NPY, FAM46A, STAC, LINC01014, ANAPC1P1, OTOF, DSP, AOX1, SST, CHODL
Sst_1	Sst+ GABAergic neuron_1	GABAergic neuron	EDNRA, QRFPR, LINC01324, NELL1, PAWR, CDH9, RCAN3, GRIN3A, FLT3, SST
Sst_2	Sst+ GABAergic neuron_2	GABAergic neuron	FAM89A, EPHA6, TPH2, HS6ST3, RCAN3, HTR4, FLT3, VWC2, NMU, SST
Sst_3	Sst+ GABAergic neuron_3	GABAergic neuron	GRIK1, NR2F2-AS1, NMU, THSD7A, EPSTI1, ZNF385D, ELFN1, ZMAT4, DKK2, SST
Sst_4	Sst+ GABAergic neuron_4	GABAergic neuron	NMU, TMTC2, KLHL14, PTPRD, LINC00943, GRID2, C8orf87, MPPED2, PDCL2, SST
Sst_5	Sst+ GABAergic neuron_5	GABAergic neuron	HGF, SEMA5A, ADGRF2, ERBB4, ITGA1, ZNF536, RND3, NEGR1, NXPH1, SST
Sst_6	Sst+ GABAergic neuron_6	GABAergic neuron	KLHL14, L3MBTL4, ADAMTS20, PREX2, FBN2, SLC8A1, DCN, C8orf34, MCAM, SST
Sst_7	Sst+ GABAergic neuron_7	GABAergic neuron	TH, SEMA6A, GABRQ, CACNA2D1, LAYN, TRHDE, SLA, RALYL, PTPRC, SST
Vip_1	Vip+ GABAergic neuron_1	GABAergic neuron	SHISA8, ZMAT4, GPR149, KCNIP4, ASIC4, RYR3, RGS16, KLF12, CBLN1, VIP
Vip_2	Vip+ GABAergic neuron_2	GABAergic neuron	VIP, GRIN2A, PPP1R1C, CDH20, CHRNA2, NRXN1, CALB2, DCC, NPR3, TAC3
Vip_3	Vip+ GABAergic neuron_3	GABAergic neuron	SMOC1, TMEM132D, SLC22A3, EGFEM1P, CHRNA2, FRAS1, TAC3, SLIT2, PPP1R1C, VIP
Vip_4	Vip+ GABAergic neuron_4	GABAergic neuron	PTGFR, CSGALNACT1, PCDH18, PEX5L, CALB2, ADGRL3, LINC01497, KCNC2, PENK, VIP
VLMC	vascular lepotomeningeal cell	brain pericyte	COLEC12, COL1A2, TBX18, ITIH5, COL6A2, SRPX2, C7, FMO2, MFAP4, FOXD1